Porro, L. B., Rayfield, E. J., Clack, J. A., May, 2023, Computed tomography and three-dimensional reconstruction of the skull of the stem tetrapod Crassigyrinus scoticus, Watson, 1929, Journal of Vertebrate Paleontology, Volume 42, Issue 3 (2022), published online 02 May, 2023, https://doi.org/10.1080/02724634.2023.2183134
Molnar, J. L., Hutchinson, J. R., Diogo, R., Clack, J. A. & Pearce, S. E., January, 2021, Evolution of forelimb musculoskeletal function across the fish-to-tetrapod transition, Science Advances: Vol. 7, No. 4, https://advances.sciencemag.org/content/7/4/eabd7457
Dickson, B. V., Clack, J.A., Smithson, T. R. & Pierce, S. E., November 2020, Functional adaptive landscapes predict terrestrial capacity at the origin of limbs, Nature volume 589, pages 242–245(2021) https://www.nature.com/articles/s41586-020-2974-5
Johanson, Z., Jeffery, J., Challands, T., Pierce, S. E. & Clack, J. A., October 2020, A New Look at Carboniferous Rhizodontid Humeri (Sarcopterygii; Tetrapodomorpha), Journal of Vertebrate Paleontology Volume 40, 2020 - Issue 3 https://www.tandfonline.com/doi/abs/10.1080/02724634.2020.1813150
Clack, J. A. & Smithson, T. R., September 2020, A new large embolomere from East Kirkton, Scottish Journal of Geology, 56, 153-158, https://doi.org/10.1144/sjg2020-008
Ruta, M., Clack, J. A. & Smithson, T. R., August 2020, A review of the stem amniote Eldeceeon rolfei from the Viséan of East Kirkton, Scotland, Transactions of the Royal Society of Edinburgh, 111, 173-192 https://doi.org/10.1017/S1755691020000079
Ahlberg, P. E. & Clack, J. A., April 2020, The smallest known Devonian tetrapod shows unexpectedly derived features, Royal Society Open Science, https://doi.org/10.1098/rsos.192117
Clack, J. A., Smithson, T. R. & Ruta, M. April 2022, A Mississippian (early Carboniferous) tetrapod showing early diversification of the hind limbs, Communications Biology, https://doi.org/10.1038/s42003-022-03199-x
A legacy in fossils: a tribute to Stan(ley) Wood – Preface. Fraser, N. C., Smithson, T. R. and Clack, J. A. 2018. Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 108, 1-5.
Clack, J. A. 2017 The East Kirkton Lagerstätte: A window onto Early Carboniferous land ecosystems. In N. C Fraser and H. D. Sues (Eds) Terrestrial Lagerstätten. Dunedin. Chapter 2, 39-64.
Clack, J.A., Fay R. & Popper A. N. (Eds) 2016 Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Acoustical Research Volume 59.
Clack, J.A. 2016. Vertebrate Diversity in a Sensory System: The Fossil Record of Otic Evolution. In Clack, J.A., Fay R. & Popper A. N. (Eds) Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Acoustical Research Volume 59 pp1-16
Clack J. A., Ahlberg, P. E. 2016. Sarcopterygians: From Lobe-Finned Fishes to the Tetrapod Stem Group. In Clack, J.A., Fay R. & Popper A. N. (Eds) Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Acoustical Research Volume 59 pp 51-70
Clack J. A. & Anderson J. S. 2016. Early Tetrapods: Experimenting with Form and Function. In Clack, J.A., Fay R. & Popper A. N. (Eds) Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Acoustical Research Volume 59 pp 71-106
Clack JA. 2016 Land vertebrates, the origin and evolution of. Encyclopedia of Evolutionary Biology, vol. 2, (ed. Kilman RM), pp. 296-304. Oxford: Academic Press.
Clack, J. A. and Milner, A. R. 2015 Early tetrapods. Handbook of Paleoherpetology. Part 3A1. Friedrich Pfeil, Munich.
Clack, J. A. 2012 Gaining Ground: the origin and evolution of tetrapods, Second Edition. Indiana University Press , Bloomington, Indiana, USA. 544pp ISBN 978-0-253-35675-8
Clack, J. A. 2000 Gaining Ground: the origin and evolution of tetrapods. (In Japanese). Kodansha, Japan.
Beeby, E. L., Clack, J. A., & Smithson, T. R. 2020. Systematics and description of the lungfish Sagenodus from the Carboniferous of the United Kingdom. Environmental Science Transactions of the Royal Society of Edinburgh. https://doi.org/10.1017/S1755691019000203.
Herbst, E.D., Doube, M., Smithson, T. R., Clack, J.A. and Hutchinson, J. R. 2019. Bony lesions in early tetrapods and the evolution of mineralized tissue repair. Paleobiology 1–22 DOI: 10.1017/pab.2019.31.
Clack, J. A., Milner A. R., Ruta, M., Marshall, J. E .A., Smithson, T. R., Smithson, K.Z. 2019. The earliest heterodont and durophagous tetrapod. RS Open Science. https://dx.doi.org/10.1098/rsos.182087
Challands, T. J., Smithson T. R., Clack, J. A., Bennett, C. E., Marshall, J. E. A., Finney, S. M., Hill, H. A Lungfish survivor of the end-Devonian extinction and an Early Carboniferous dipnoan radiation. Journal of Systematic Palaeontology. https://doi.org/10.1080/14772019.2019.1572234.
Clack, J.A., Smithson T.R., Bennett, C.E., Davies, S.J., Scott A.C. 2019. A Tournaisian (earliest Carboniferous) conglomerate-preserved non-marine faunal assemblage and its environmental and sedimentological context. PeerJ https://doi.org/10.7717/peerj.5972.
Clack, J. A., Challands, T. J., Smithson, T. R., and Smithson, K. Z. 2018b Newly recognized Famennian lungfishes from East Greenland reveal dental diversity and blur the Devonian-Carboniferous boundary. Papers in Palaeontology doi: 10.1002/spp2.1242
Clack J. A., Porro L.B. and Bennett, C.E. 2018a. A Crassigyrinus-like jaw from the Tournaisian (Early Mississippian) of Scotland. Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 108, 37-46
Otoo, B. K. A., Clack, J. A., Smithson, T. R., Bennett, C. E., Kearsey, T. I., Coates, M. I. 2018. An early post-Devonian vertebrate fauna preserved in Scottish Tournaisian flood plain deposits. Palaeontology. doi: 10.1111/pala.12395
Smithson, T. R and Clack, J. A. 2018. A new tetrapod from Romer’s Gap reveals an early adaptation for walking. Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 108, 89-97
Richards K.R., J.E. Sherwin, T.R. Smithson, R.F. Bennion, S.J. Davies, J.E. A.Marshall, and J.A. Clack. 2018. Diverse and durophagous: Early Carboniferous chondrichthyans from the Scottish Borders. Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 108, 1-21.
Smithson, T. R., Brown, A. E., Davies, S. J., Marshall, J. E. A., Millward, D., Walsh, S. A. and Clack, J. A. 2017 A new Mississippian tetrapod from Fife, Scotland, and its environmental context. Papers in Palaeontology 2017, 1-11. DOI: 10.1002/spp2.1086
Bennett, C., Kearsey, T., Davies, S., Millward, D., Clack, J., Smithson, T. and Marshall, J. 2016 Early Carboniferous sandy siltstones deposited in seasonal flooding episodes preserve rare non-marine fossils. Sedimentology 63, 1677-1700 doi: 10.1111/sed.12280
Sanchez, S., Tafforeau, P., Clack, J. A. and Ahlberg, P. E. 2016 Life history of the stem tetrapod Acanthostega revealed by synchrotron microtomography. Nature. 537, 408-411. Doi.10.1038/nature19354.
Clack, J. A., Bennett, C. E., Carpenter, D. K., Davies, S. J., Fraser, N. C., Kearsey, T. I., Marshall, J. E. A., Millward, D., Otoo, B. K. A., Reeves, E. J., Ross, A. J., Ruta, M., Smithson, K. Z., Smithson, T. R. and Walsh, S. 2016 Phylogenetic and Environmental Context of a Tournaisian Tetrapod Fauna. Nature Ecology and Evolution. doi:10.1038/s41559-016-0002 Published online December 2016, in print January 2017
Smithson, T. R., Richards, K. R. and Clack, J. A. 2015 Lungfish diversity in Romer’s Gap: reaction to the end-Devonian extinction. Palaeontology doi: 10.1111/pala.12203
Clack, J. A. and Milner, A. R. 2015 Basal tetrapods. Handbook of Paleoherpetology. Part 3A1. Friedrich Pfeil, Munich.
Porro, L.B., Rayfield, E. M., and Clack, J. A. 2015 Computed tomography, anatomical description and three-dimensional reconstruction of the lower jaw of Eusthenopteron foordi Whiteaves ,1881 from the Upper Devonian of Canada. Palaeontology 2015, 1-17
Anderson, J. S., Smithson, T. R., Mansky, C., Meyer, T., Carroll, R. L. and Clack, J. A. 2015 A diverse tetrapod fauna at the base of Romer’s Gap. PLoS ONE 10(4): e0125446. doi:10.1371/journal.pone.0125446
Porro, L. B., Rayfield, E. J. and Clack, J. A. 2015 Descriptive anatomy and three-dimensional reconstruction of the skull of the early tetrapod Acanthostega gunnari Jarvik, 1952. PLoS ONE
Sookias, R. B, Böhmer, C. and Clack, J. A. 2014 Redescription and phylogenetic analysis of the mandible of an enigmatic Late Carboniferous tetrapod from Nova Scotia and the lability of Meckelian jaw ossification. PLoS One. October 2014, 9, e109717
Klembara, J., Clack, J. A., Milner, A. R., Ruta, M. 2014 Cranial anatomy, ontogeny, and relationships of the Late Carboniferous tetrapod Gephyrostegus bohemicus Jaekel, 1902. Journal of Vertebrate Paleontology. 34, 774-792.
Smithson, T. R. and Clack, J. A. 2013 Tetrapod appendicular skeletal elements from the Early Carboniferous of Scotland. Comptes Rendus. doi.org/10.1016/j.crpv.2013.06.006.
Pierce, S. E, Hutchinson, J. R. and Clack, J. A. 2013 Historical perspectives on the evolution of tetrapodomorph movement. Integrative and Comparative Biology. Integrative & Comparative Biology pp. 1-15.
Pierce, S. E., Ahlberg, P. E., Hutchinson, J. R., Molnar, J. L., Sanchez, S., Tafforeau, P. and Clack, J. A. 2013. Vertebral architecture in the earliest stem tetrapods. Nature 494:226-230.
Clack, J. A., Witzmann, F., Snyder D., and Müller, J. A. 2012 colosteid-like early tetrapod from the St. Louis Limestone (Early Carboniferous, Meramecian), St. Louis, Missouri, USA. Fieldiana Life and Earth Sciences 5, 17-39.
Sharp, E. L. and Clack, J. A. 2012 Redescription of the lungfish Straitonia waterstoni from the Viséan of Lothian, Scotland. Transactions of the Royal Society of Edinburgh, Earth and Environmental Sciences 102, 179-189.
Pierce, S. E., Clack, J. A. and Hutchinson, J. R. 2012 Three-dimensional limb joint mobility in the early tetrapod Ichthyostega. Nature, 486:523-526.
Smithson, T.R., Wood, S.P., Marshall, J.E.A. and Clack, J.A. 2012 Earliest Carboniferous tetrapod and arthropod faunas from Scotland populate Romer's Gap. Proceedings of the National Academy of Sciences. 109:4532-4537
Clack, J. A., Ahlberg, P. E., Blom, H., and Finney, S. M. 2012 A new genus of Devonian tetrapod from East Greenland with new information on the lower jaw of Ichthyostega. Palaeontology 55: 73-86
Molnar, J., Pierce, S. E., Clack, J. A. and Hutchinson, J. R. 2012 Idealized landmark-based geometric reconstructions of poorly preserved fossil material: a case study of an early tetrapod vertebra. Paleontologica Electronica. 15:1.32
Clack, J. A. 2011 A new microsaur from the Early Carboniferous (Viséan) of East Kirkton, Scotland, showing soft tissue evidence. Special Papers in Palaeontology 86:45-56.
Clack, J. A. 2011 A Carboniferous embolomere tail with supraneural radials. Journal of Vertebrate Paleontology. 31:1150-1153
Polgar, G., Malavasi, S., Cipolato, G., Georgalas, V., Clack, J. A. and Torricelli, P. 2011 Acoustic communication at the water's edge: Evolutionary insights from a mudskipper. PLoS ONE 6(6): e21434
Pierce, S. E., Clack J. A. and Hutchinson J. R. 2011 Comparative axial morphology in pinnipeds and its correlation with aquatic locomotory behaviour. Journal of Anatomy 219: 502-514
Clack. J. A. and Milner, A. R. 2019 The Morphology and Systematics of the Pennsylvanian Amphibian Platyrhinops lyelli (Amphibia: Temnospondyli). Transactions of the Royal Society of Edinburgh, Earth and Environmental Sciences 100, 275-295.
Klembara, J., Clack, J. A. and Cernansky, A. 2010 The anatomy of the palate of Chroniosaurus dongusensis (Chroniosuchia, Chroniosuchidae) from the Upper Permian of Russia. Palaeontology, 53: 1147-1153.
Callier, V., Clack, J. A., Ahlberg, P. E. 2009 Contrasting ontogenetic trajectories in the earliest known tetrapod forelimbs. Science. 324, 364-367. DOI: 10.1126/science. 1167542
Clack, J. A. and Klembara J. 2009 An articulated specimen of a chroniosuchid from the Late Permian of Russia. Special Papers in Palaeontology. 81: 15-42.
Clack, J. A. 2009 The fish-tetrapod transition: new fossils and interpretations. Evolution: Education and Outreach. 2: 213-223
Clack, J. A. 2009 The fin to limb transition: new data, interpretations and hypotheses from paleontology and developmental biology. Annual Review of Earth and Planetary Sciences. 37: 163-179
Daeschler, E. B, Clack, J. A., Shubin, N. H. 2009 Late Devonian tetrapod remains from Red Hill Pennsylvania, USA: how much diversity? Ahlberg, P. E., Blom, H. & Boisvert, C. A. (eds.) Forty years of early vertebrates: papers from the 11th International Symposium on Early and Lower Vertebrates. Acta Zoologica Special Issue, Supplement to Vol. 90.306-317.
Kimmel, C, Sidlauskas, B. and Clack, J. A. 2009 Linked morphological changes during palate evolution in early tetrapods. Journal of Anatomy 215: 91-109 doi: 10.1111/j.1469-7580.2009.01108.
Ahlberg, P.E., Clack, J. A., Luksevics, E., Blom, H. and Zupins I. 2008 Ventastega curonica and the origin of tetrapod morphology. Nature 453, 1199-1204
Larsen, H., Olsen,P., and Clack, J. A. 2008. The Devonian Basin in East Greenland - Review of basin evolution and vertebrate assemblages. In The Greenland Caledonides: Evolution of the Northeast Margin of Laurentia. eds A. K. Higgins, J. A.Gilotti and M. P. Smith. Geological Society of America Memoir. pp. 273-292. doi 10.1130/2008.1202(11).
Blom, H., Clack, J. A., Ahlberg P. E. and Friedman, M. 2007 Devonian vertebrates from East Greenland: a review of faunal composition and distribution. Geodiversitas: 29: 119-141
Clack, J. A. 2007 Devonian climate change, breathing and the origin of the tetrapod stem group. Integrative and Comparative Biology. 47: 510-523.
Clack, J. A. 2006 The emergence of early tetrapods. Palaeogeography, Palaeoclimatology,Palaeoecology. 232, 167-189
Ruta, M. and Clack, J. A. 2006 A review of Silvanerpeton miripedes, a stem amniote from the Lower Carboniferous of East Kirkton, West Lothian, Scotland. Trans. R. Soc. Ed.: Earth Sciences, 97, 31-63
Ahlberg, P. E., Clack, J. A. and Blom, H. 2005 The axial skeleton of the Devonian tetrapod Ichthyostega. Nature. 437, 137-140.
Blom, H., Clack, J. A. & Ahlberg, P. E. 2005. Localities, distribution and stratigraphical context of the Late Devonian tetrapods of East Greenland. Meddelelser om Grønland, Geosciences. 43, 1-50.
Clack, J. A. 2005 Making headway and finding a foothold: tetrapods come ashore. In: Evolving form and function: fossils and development. D. E. G. Briggs, (ed.) Peabody Museum of Natural History Special Publications, Yale University. pp 223-244.
Clack, J. A. and Finney, S. M. 2005 Pederpes finneyae, an articulated tetrapod from the Tournaisian of Western Scotland. Journal of Systematic Palaeontology. 2: 311-346.
Clack, J. A. and Ahlberg, P. E. 2004 A new stem tetrapod from the Early Carboniferous of Northern Ireland. In Recent advances in the origin and early radiation of vertebrates. Arratia, G., Cloutier, R., Wilson, M. V. H. (eds) Verlag Friedrich Pfeil pp. 309-320
Clement, G., Ahlberg, P. E., Blieck, A., Blom, H., Clack, J. A., Poty, E., Thorez, J., & Janvier, P. 2004 Devonian tetrapod from western Europe. Nature 427: 412-413 (Brief Comm.)
Clack, J. A., Ahlberg P. E., Finney, S. M, Dominguez Alonso P., Robinson, J and Ketcham, R. A. 2003. A uniquely specialised ear in a very early tetrapod. Nature. 425, 65-69
Clack, J. A. 2003 A revised reconstruction of the dermal skull roof of Acanthostega, an early tetrapod from the Late Devonian. Trans R. Soc. Ed. Earth Sciences: 93, 163-165
Clack, J. A. 2003. A new baphetid (stem tetrapod) from the Upper Carboniferous of Tyne and Wear, UK, and the evolution of the tetrapod occiput. Can. J. Earth Sci. 40: 483-498
Clack, J. A. 2002 The dermal skull roof of Acanthostega gunnari, an early tetrapod from the Late Devonian. Trans. R. Soc. Ed. 93: 17-33.
Clack, J. A. 2002 Patterns and processes in the early evolution of the tetrapod ear. Journal of Neurobiology 53, 251-264
Clack, J. A. 2002 An early tetrapod from 'Romers Gap'. Nature 418: 72-76.
Clack J. A. and Anderson J. S. 2016 Early Tetrapods: Experimenting with Form and Function. In Clack, J.A., Fay R. and Popper A. N. (Eds) Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Auditory Research Volume 59 pp 71-106
Clack J. A. and Ahlberg, P. E. 2016 Sarcopterygians: From Lobe-Finned Fishes to the Tetrapod Stem Group. In Clack, J.A., Fay R. and Popper A. N. (Eds) Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Auditory Research Volume 59 pp 51-70
Clack, J.A. 2016 Vertebrate Diversity in a Sensory System: The Fossil Record of Otic Evolution. In Clack, J.A., Fay R. and Popper A. N. (Eds) Evolution of the Vertebrate Ear: Evidence from the Fossil Record. Springer Handbook of Auditory Research Volume 59 pp 1-16
Clack, J. A. 2013 Fossil vertebrates - Palaeozoic non-amniote tetrapods. Earth Systems and Environmental Science. Elsevier (Online update of Encyclopaedia of Geology 2005. Elsevier, Oxford 2: 468-497)
Clack, J. A., Sharp, E. L. and Long, J. A. 2011 The fossil record of lungfishes. In The Biology of Lungfishes, J. M. Jørgensen and Joss, J. eds. Science Publishers Inc. Enfield, New Hampshire, USA pp. 1-42.
Clack, J. A. 2005 Fossil vertebrates: Palaeozoic non-amniote tetrapods. Encyclopaedia of Geology. Elsevier, Oxford 2: 468-497
Clack, J. A., Allin, E. 2004 The evolution of single- and multiple-ossicle ears in fishes and tetrapods. In: Evolution of the Vertebrate Auditory System, Manley, G. A., Popper, A. and Fay, R. R. (eds.) New York, Springer-Verlag. pp 238-163
Manley, G. A., Clack, J. A. 2004 An outline of the evolution of vertebrate hearing organs. In: Evolution of the Vertebrate Auditory System, Manley, G. A., Popper, A. and Fay, R. R. (eds.) New York, Springer-Verlag. pp 1-26
Clack, J. A. 2000 The origin of tetrapods. Amphibian Biology Volume 4, Paleontology; Chapter 2 Heatwole, H. and Carroll, R. L. (eds.) Surrey Beatty pp. 973-1029
Clack, J. A. and Carroll, R. L. 2000 Early Carboniferous tetrapods. Amphibian Bology Volume 4, Paleontology; Chapter 2 Heatwole, H. and Carroll, R. L. (eds.) Surrey Beatty pp. 1030-1043
Clack, J. A. 1997 World Wide Web pages for University of Arizona 'Tree of Life' describing the taxa Acanthostega, Ichthyostega, Crassigyrinus and the baphetid tetrapods. Updated 2006.
Ahlberg, P. E. and Clack, J. A. 2006 A firm step from water to land. News and Views, Nature 440, 747-749
Clack, J. A. 2004 From Fins to Fingers, Perspectives; Science, 304, 57-58
Ruta, M., Clack, J. A. and Milner, A. C. (eds.) 2009 Patterns and processes in early vertebrate evolution. Special Papers in Palaeontology 81: 1-173.
Sánchez-Villagra, M. R. and Clack, J. A. (eds.) 2004 Fossils of the Miocene Castillo Formation, Venezuela: Contributions in Neotropical Palaeontology. Special Papers in Palaeontology 71: 1-112.
Clack, J. A. 2010 What is a tetrapod? Grzimek's Animal Life. Gale Web 17 May.
Clack, J. A. 2008 Overview from the chair: Sea to Sand. Proceedings of the 2007 Moray Society Conference. Moray Society, Elgin Museum, Elgin, Moray, Scotland. pp. 7-10
Clack, J. A. 2008 Devonian Missing Link: Tiktaalik roseae. McGraw-Hill Yearbook of Science & Technology, 10th Edition, McGraw-Hill, New York, 2008, pp. 91-93.
Clack, J. A. 2007 Anthracosauria, In: McGraw-Hill Encyclopedia of Science & Technology, Volume 2, McGraw-Hill, New York, 2007, pp. 30-31.
Clack, J. A. 2006 From fins to limbs. Natural History Magazine, 115, 36-41
Clack, J. A. 2006 Tetrapod origins. McGraw-Hill Yearbook of Science & Technology, McGraw-Hill, New York, 2006, pp. 325-327.
Clack, J. A. 2006 Le premier pied à terre. Pour la Science. 340, 30-36 (Translation of Scientific American article below.)
Clack, J. A. 2005 Getting a leg up on land. Invited article. Scientific American. December, 293, 100-107
Last updated 6th June, 2019 by Rob Clack